You did not define your gateway (192.168.1.1) for interface vmbr1. I may be wrong, but this would be what I would try first.
You have...
auto vmbr1
iface vmbr1 inet static
address 192.168.1.31
netmask 255.255.255.0
bridge-ports eno2
bridge-stp off
bridge-fd 0
...when it...